as any wild rock-pigeon! We can understand these facts, on the
well-known principle of reversion to ancestral characters, if all the
domestic breeds have descended from the rock-pigeon. But if we deny
this, we must make one of the two following highly improbable
suppositions. Either, firstly, that all the several imagined
aboriginal stocks were coloured and marked like the rock-pigeon,
although no other existing species is thus coloured and marked, so
that in each separate breed there might be a tendency to revert to the
very same colours and markings. Or, secondly, that each breed, even
the purest, has within a dozen or, at most, within a score of
generations, been crossed by the rock-pigeon: I say within a dozen or
twenty generations, for we know of no fact countenancing the belief
that the child ever reverts to some one ancestor, removed by a greater
number of generations. In a breed which has been crossed only once
with some distinct breed, the tendency to reversion to any character
derived from such cross will naturally become less and less, as in
each succeeding generation there will be less of the foreign blood;
but when there has been no cross with a distinct breed, and there is a
tendency in both parents to revert to a character, which has been lost
during some former generation, this tendency, for all that we can see
to the contrary, may be transmitted undiminished for an indefinite
number of generations. These two distinct cases are often confounded
in treatises on inheritance.

Lastly, the hybrids or mongrels from between all the domestic breeds
of pigeons are perfectly fertile. I can state this from my own
observations, purposely made on the most distinct breeds. Now, it is
difficult, perhaps impossible, to bring forward one case of the hybrid
offspring of two animals CLEARLY DISTINCT being themselves perfectly
fertile. Some authors believe that long-continued domestication