Darwin and Modern Science by Sir Albert Charles Seward
page 91 of 912 (09%)
page 91 of 912 (09%)
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of plants had all disappeared in eleven days.
The experiments of Poulton and Sanders ("Report of the British Association" (Bristol, 1898), London, 1899, pages 906-909.) were made with 600 pupae of Vanessa urticae, the "tortoise-shell butterfly." The pupae were artificially attached to nettles, tree-trunks, fences, walls, and to the ground, some at Oxford, some at St Helens in the Isle of Wight. In the course of a month 93 per cent of the pupae at Oxford were killed, chiefly by small birds, while at St Helens 68 per cent perished. The experiments showed very clearly that the colour and character of the surface on which the pupa rests--and thus its own conspicuousness--are of the greatest importance. At Oxford only the four pupae which were fastened to nettles emerged; all the rest--on bark, stones and the like--perished. At St Helens the elimination was as follows: on fences where the pupae were conspicuous, 92 per cent; on bark, 66 per cent; on walls, 54 per cent; and among nettles, 57 per cent. These interesting experiments confirm our views as to protective coloration, and show further, THAT THE RATIO OF ELIMINATION IN THE SPECIES IS A VERY HIGH ONE, AND THAT THEREFORE SELECTION MUST BE VERY KEEN. We may say that the process of selection follows as a logical necessity from the fulfilment of the three preliminary postulates of the theory: variability, heredity, and the struggle for existence, with its enormous ratio of elimination in all species. To this we must add a fourth factor, the INTENSIFICATION of variations which Darwin established as a fact, and which we are now able to account for theoretically on the basis of germinal selection. It may be objected that there is considerable uncertainty about this LOGICAL proof, because of our inability to demonstrate the selection- value of the initial stages and the individual stages of increase. We have therefore to fall back on PRESUMPTIVE EVIDENCE. This is to be found in THE |
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