The Movements and Habits of Climbing Plants by Charles Darwin
page 58 of 178 (32%)
page 58 of 178 (32%)
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petioles into contact with some supporting object. The petiole of a
young leaf, after a slight rub, became curved in 6 m.; another, on a cold day, in 20 m., and others in from 8 m. to 10 m. Their curvature usually increased greatly in from 15 m. to 20 m., and they became straight again in between 5 hrs. and 6 hrs., but on one occasion in 3 hrs. When a petiole has fairly clasped a stick, it is not able, on the removal of the stick, to straighten itself. The free upper part of one, the base of which had already clasped a stick, still retained the power of movement. A loop of thread weighing 0.125th of a grain caused a petiole to curve; but the stimulus was not sufficient, the loop remaining suspended, to cause a permanent flexure. If a much heavier loop be placed in the angle between the petiole and the stem, it produces no effect; whereas we have seen with Clematis montana that the angle between the stem and petiole is sensitive. Tropaeolum peregrinum.--The first-formed internodes of a young plant did not revolve, resembling in this respect those of a twining plant. In an older plant the four upper internodes made three irregular revolutions, in a course opposed to the sun, at an average rate of 1 hr. 48 min. It is remarkable that the average rate of revolution (taken, however, but from few observations) is very nearly the same in this and the two last species, namely, 1 hr. 47 m., 1 hr. 46 m., and 1 hr. 48 m. The present species cannot twine spirally, which seems mainly due to the rigidity of the stem. In a very young plant, which did not revolve, the petioles were not sensitive. In older plants the petioles of quite young leaves, and of leaves as much as an inch and a quarter in diameter, are sensitive. A moderate rub caused one to curve in 10 m., and others in 20 m. They became straight again in between 5 hrs. 45m. and 8 hrs. Petioles which have naturally come into contact with a stick, sometimes take two turns |
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