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Variation of Animals and Plants under Domestication, the — Volume 1 by Charles Darwin
page 132 of 624 (21%)
than once met with Cagias (a sub-Himalayan domestic race) possessed of four
teats." (3/80. 'Journal Asiat. Soc. of Bengal' volume 16 1847 page 1015.)
This case is the more remarkable as, when any part or organ is present in
reduced number in comparison with the same part in allied groups, it
usually is subject to little variation. The presence of interdigital pits
has likewise been considered as a generic distinction in sheep; but Isidore
Geoffroy (3/81. 'Hist. Nat. Gen.' tome 3 page 435.) has shown that these
pits or pouches are absent in some breeds.

In sheep there is a strong tendency for characters, which have apparently
been acquired under domestication, to become attached either exclusively to
the male sex, or to be more highly developed in this than in the other sex.
Thus in many breeds the horns are deficient in the ewe, though this
likewise occurs occasionally with the female of the wild musmon. In the
rams of the Wallachian breed, "the horns spring almost perpendicularly from
the frontal bone, and then take a beautiful spiral form; in the ewes they
protrude nearly at right angles from the head, and then become twisted in a
singular manner." (3/82. 'Youatt on Sheep' page 138.) Mr. Hodgson states
that the extraordinarily arched nose or chaffron, which is so highly
developed in several foreign breeds, is characteristic of the ram alone,
and apparently is the result of domestication. (3/83. 'Journal Asiat. Soc.
of Bengal' volume 16 1847 pages 1015, 1016.) I hear from Mr. Blyth that the
accumulation of fat in the fat-tailed sheep of the plains of India is
greater in the male than in the female; and Fitzinger (3/84. 'Racen des
Zahmen Schafes' s. 77.) remarks that the mane in the African maned race is
far more developed in the ram than in the ewe.

Different races of sheep, like cattle, present constitutional differences.
Thus the improved breeds arrive at maturity at an early age, as has been
well shown by Mr. Simonds through their early average period of dentition.
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