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Variation of Animals and Plants under Domestication, the — Volume 1 by Charles Darwin
page 59 of 624 (09%)
and one to nearly four. In the Scotch deer-hound there is a striking and
remarkable difference in the size of the male and female. (1/64. W. Scrope
'Art of Deer-Stalking' page 354.) Every one knows how the ears vary in size
in different breeds, and with their great development their muscles become
atrophied. Certain breeds of dogs are described as having a deep furrow
between the nostrils and lips. The caudal vertebrae, according to F.
Cuvier, on whose authority the two last statements rest, vary in number;
and the tail in English cattle and some shepherd dogs is almost absent. The
mammae vary from seven to ten in number; Daubenton, having examined twenty-
one dogs, found eight with five mammae on each side; eight with four on
each side; and the others with an unequal number on the two sides. (1/65.
Quoted by Col. Ham. Smith in 'Nat. Lib.' volume 10 page 79.) Dogs have
properly five toes in front and four behind, but a fifth toe is often
added; and F. Cuvier states that, when a fifth toe is present, a fourth
cuneiform bone is developed; and, in this case, sometimes the great
cuneiform bone is raised, and gives on its inner side a large articular
surface to the astragalus; so that even the relative connection of the
bones, the most constant of all characters, varies. These modifications,
however, in the feet of dogs are not important, because they ought to be
ranked, as De Blainville has shown (1/66. De Blainville 'Osteographie,
Canidae' page 134. F. Cuvier 'Annales du Museum' tome 18 page 342. In
regard to mastiffs, see Col. H. Smith 'Nat. Lib.' volume 10 page 218. For
the Thibet mastiff, see Mr. Hodgson in 'Journal of As. Soc. of Bengal'
volume 1 1832 page 342.) as monstrosities. Nevertheless they are
interesting from being correlated with the size of the body, for they occur
much more frequently with mastiffs and other large breeds than with small
dogs. Closely allied varieties, however, sometimes differ in this respect;
thus Mr. Hodgson states that the black-and-tan Lassa variety of the Thibet
mastiff has the fifth digit, whilst the Mustang sub-variety is not thus
characterised. The extent to which the skin is developed between the toes
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