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Variation of Animals and Plants under Domestication, the — Volume 2 by Charles Darwin
page 105 of 776 (13%)
good case: but Mr. Wilkinson states (16/10. Mr. J. Wilkinson in 'Remarks
addressed to Sir J. Sebright' 1820 page 38.), that a breed derived from this
same cross was actually established in another part of England; and if it had
failed in fertility, the fact would surely have been noticed. Moreover,
supposing that Mr. Youatt had proved his case, it might be argued that the
sterility was wholly due to the two parent-breeds being descended from
primordially distinct species.

In the case of plants Gartner states that he fertilised thirteen heads (and
subsequently nine others) on a dwarf maize bearing yellow seed (16/11.
'Bastarderzeugung' s. 87, 169. See also the Table at the end of volume.) with
pollen of a tall maize having red seed; and one head alone produced good seed,
but only five in number. Though these plants are monoecious, and therefore do
not require castration, yet I should have suspected some accident in the
manipulation, had not Gartner expressly stated that he had during many years
grown these two varieties together, and they did not spontaneously cross; and
this, considering that the plants are monoecious and abound with pollen, and
are well known generally to cross freely, seems explicable only on the belief
that these two varieties are in some degree mutually infertile. The hybrid
plants raised from the above five seeds were intermediate in structure,
extremely variable, and perfectly fertile. (16/12. 'Bastarderzeugung' s. 87,
577.) In like manner Prof. Hildebrand (16/13. 'Bot. Zeitung' 1868 page 327.)
could not succeed in fertilising the female flowers of a plant bearing brown
grains with pollen from a certain kind bearing yellow grains; although other
flowers on the same plant, which were fertilised with their own pollen,
yielded good seed. No one, I believe, even suspects that these varieties of
maize are distinct species; but had the hybrids been in the least sterile, no
doubt Gartner would at once have so classed them. I may here remark, that with
undoubted species there is not necessarily any close relation between the
sterility of a first cross and that of the hybrid offspring. Some species can
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