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Variation of Animals and Plants under Domestication, the — Volume 2 by Charles Darwin
page 29 of 776 (03%)
The Equus indicus (13/36. Sclater in 'Proc. Zoolog. Soc.' 1862 page 163: this
species is the Ghor-Khur of N.W. India, and has often been called the Hemionus
of Pallas. See also Mr. Blyth's excellent paper in 'Journal of Asiatic Soc. of
Bengal' volume 28 1860 page 229.) is characterised by a spinal stripe, without
shoulder or leg stripes; but traces of these latter stripes may occasionally
be seen even in the adult (13/37. Another species of wild ass, the true E.
hemionus or Kiang, which ordinarily has no shoulder-stripes, is said
occasionally to have them; and these, as with the horse and ass, are sometimes
double: see Mr. Blyth in the paper just quoted and in 'Indian Sporting Review'
1856 page 320: and Col. Hamilton Smith in 'Nat. Library, Horses' page 318; and
'Dict. Class. d'Hist. Nat.' tome 3 page 563.) and Colonel S. Poole, who has
had ample opportunities for observation, informs me that in the foal, when
first born, the head and legs are often striped, but the shoulder-stripe is
not so distinct as in the domestic ass; all these stripes, excepting that
along the spine, soon disappear. Now a hybrid, raised at Knowsley (13/38.
Figured in the 'Gleanings from the Knowsley Menageries' by Dr. J.E. Gray.)
from a female of this species by a male domestic ass, had all four legs
transversely and conspicuously striped, had three short stripes on each
shoulder and had even some zebra-like stripes on its face! Dr. Gray informs me
that he has seen a second hybrid of the same parentage, similarly striped.

From these facts we see that the crossing of the several equine species tends
in a marked manner to cause stripes to appear on various parts of the body,
especially on the legs. As we do not know whether the parent-form of the genus
was striped, the appearance of the stripes can only hypothetically be
attributed to reversion. But most persons, after considering the many
undoubted cases of variously coloured marks reappearing by reversion in my
experiments on crossed pigeons and fowls, will come to the same conclusion
with respect to the horse-genus; and if so, we must admit that the progenitor
of the group was striped on the legs, shoulders, face, and probably over the
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