Variation of Animals and Plants under Domestication, the — Volume 2 by Charles Darwin
page 47 of 776 (06%)
page 47 of 776 (06%)
|
development, it is difficult to understand how an organ arrested at a very
early period of growth should acquire its full functional perfection;--how a petal, supposed to be thus arrested, should acquire its brilliant colours, and serve as an envelope to the flower, or a stamen produce efficient pollen; yet this occurs with many peloric flowers. That pelorism is not due to mere chance variability, but either to an arrest of development or to reversion, we may infer from an observation made by Ch. Morren (13/70. In his discussion on some curious peloric Calceolarias quoted in 'Journal of Horticulture' February 24, 1863 page 152.) namely, that families which have irregular flowers often "return by these monstrous growths to their regular form; whilst we never see a regular flower realise the structure of an irregular one." Some flowers have almost certainly become more or less completely peloric through reversion, as the following interesting case shows. Corydalis tuberosa properly has one of its two nectaries colourless, destitute of nectar, only half the size of the other, and therefore, to a certain extent, in a rudimentary state; the pistil is curved towards the perfect nectary, and the hood, formed of the inner petals, slips off the pistil and stamen in one direction alone, so that, when a bee sucks the perfect nectary, the stigma and stamens are exposed and rubbed against the insect's body. In several closely allied genera, as in Dielytra, etc., there are two perfect nectaries, the pistil is straight, and the hood slips off on either side, according as the bee sucks either nectary. Now, I have examined several flowers of Corydalis tuberosa, in which both nectaries were equally developed and contained nectar; in this we see only the redevelopment of a partially aborted organ; but with this redevelopment the pistil becomes straight, and the hood slips off in either direction, so that these flowers have acquired the perfect structure, so well adapted for insect agency, of Dielytra and its allies. We cannot attribute these coadapted modifications to chance, or to correlated variability; we must attribute them to reversion to a primordial condition of |
|