Variation of Animals and Plants under Domestication, the — Volume 2 by Charles Darwin
page 63 of 776 (08%)
page 63 of 776 (08%)
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obtained the same result from crossing a peloric Linaria with the common form.
I carefully examined the flowers of ninety plants of the crossed Antirrhinum in the two beds, and their structure had not been in the least affected by the cross, except that in a few instances the minute rudiment of the fifth stamen, which is always present, was more fully or even completely developed. It must not be supposed that this entire obliteration of the peloric structure in the crossed plants can be accounted for by any incapacity of transmission; for I raised a large bed of plants from the peloric Antirrhinum, artificially fertilised by its own pollen, and sixteen plants, which alone survived the winter, were all as perfectly peloric as the parent-plant. Here we have a good instance of the wide difference between the inheritance of a character and the power of transmitting it to crossed offspring. The crossed plants, which perfectly resembled the common snapdragon, were allowed to sow themselves, and out of a hundred and twenty-seven seedlings, eighty-eight proved to be common snapdragons, two were in an intermediate condition between the peloric and normal state, and thirty-seven were perfectly peloric, having reverted to the structure of their one grand-parent. This case seems at first sight to offer an exception to the rule just given, namely, that a character which is present in one form and latent in the other is generally transmitted with prepotent force when the two forms are crossed. For in all the Scrophulariaceae, and especially in the genera Antirrhinum and Linaria, there is, as was shown in the last chapter, a strong latent tendency to become peloric; but there is also, as we have seen, a still stronger tendency in all peloric plants to reacquire their normal irregular structure. So that we have two opposed latent tendencies in the same plants. Now, with the crossed Antirrhinums the tendency to produce normal or irregular flowers, like those of the common Snapdragon, prevailed in the first generation; whilst the tendency to pelorism, appearing to gain strength by the intermission of a generation, prevailed to a large extent in the second set of seedlings. How it is possible for a character to gain strength by the intermission of a generation, will be considered in the |
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