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Variation of Animals and Plants under Domestication, the — Volume 2 by Charles Darwin
page 97 of 776 (12%)
fowls, the young of which from the first are sometimes much diversified. As
cross-bred animals are generally of large size and vigorous, they have been
raised in great numbers for immediate consumption. But for breeding they are
found utterly useless; for though they may themselves be uniform in character,
they yield during many generations astonishingly diversified offspring. The
breeder is driven to despair, and concludes that he will never form an
intermediate race. But from the cases already given, and from others which
have been recorded, it appears that patience alone is necessary; as Mr.
Spooner remarks, "nature opposes no barrier to successful admixture; in the
course of time, by the aid of selection and careful weeding, it is practicable
to establish a new breed." After six or seven generations the hoped-for result
will in most cases be obtained; but even then an occasional reversion, or
failure to keep true, may be expected. The attempt, however, will assuredly
fail if the conditions of life be decidedly unfavourable to the characters of
either parent-breed. (15/30. See Colin 'Traite de Phys. Comp. des Animaux
Domestiques' tome 2 page 536, where this subject is well treated.)

Although the grandchildren and succeeding generations of cross-bred animals
are generally variable in an extreme degree, some curious exceptions to the
rule have been observed both with crossed races and species. Thus Boitard and
Corbie (15/31. 'Les Pigeons' page 37.) assert that from a Pouter and a Runt "a
Cavalier will appear, which we have classed amongst pigeons of pure race,
because it transmits all its qualities to its posterity." The editor of the
'Poultry Chronicle' (15/32. Volume 1 1854 page 101.) bred some bluish fowls
from a black Spanish cock and a Malay hen; and these remained true to colour
"generation after generation." The Himalayan breed of rabbits was certainly
formed by crossing two sub-varieties of the silver-grey rabbit; although it
suddenly assumed its present character, which differs much from that of either
parent-breed, yet it has ever since been easily and truly propagated. I
crossed some Labrador and Penguin ducks, and recrossed the mongrels with
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