The Evolution of Man — Volume 2 by Ernst Heinrich Philipp August Haeckel
page 34 of 417 (08%)
page 34 of 417 (08%)
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ventral side of the branchial sac runs a ciliated groove--the
hypobranchial groove which we have previously found at the same spot in the Amphioxus. The food of the Ascidia also consists of tiny organisms, infusoria, diatoms, parts of decomposed marine plants and animals; etc. These pass with the water into the gill-crate and the digestive part of the gut at the end of it, at first into an enlargement of it that represents the stomach. The adjoining small intestine usually forms a loop, bends forward, and opens by an anus (Figure 2.220 a), not directly outwards, but first into the mantle cavity; from this the excrements are ejected by a common outlet (a apostrophe) together with the used-up water and the sexual products. The outlet is sometimes called the branchial pore, and sometimes the cloaca or ejection-aperture. In many of the Ascidiae a glandular mass opens into the gut, and this represents the liver. In some there is another gland besides the liver, and this is taken to represent the kidneys. The body-cavity proper, or coeloma, which is filled with blood and encloses the hepatic gut, is very narrow in the Ascidia, as in the Amphioxus, and is here also usually confounded with the wide atrium, or peribranchial cavity, full of water. There is no trace in the fully-developed Ascidia of a chorda dorsalis, or internal axial skeleton. It is the more interesting that the young animal that emerges from the ovum HAS a chorda, and that there is a rudimentary medullary tube above it. The latter is wholly atrophied in the developed Ascidia, and looks like a small nerve-ganglion in front above the gill-crate. It corresponds to the upper "gullet-ganglion" or "primitive brain" in other vermalia. Special sense-organs are either wanting altogether or are only found in a very rudimentary form, as simple optic spots and touch-corpuscles or tentacles that surround the mouth. The muscular system is very slightly and irregularly developed. |
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