The Evolution of Man — Volume 2 by Ernst Heinrich Philipp August Haeckel
page 40 of 417 (09%)
page 40 of 417 (09%)
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segmentation-cells get more or less irregularly displaced, while the
segmentation-cavity enlarges in the centre of the morula; in the end the former all lie on the surface of the latter, so that the foetus attains the familiar blastula shape and forms a hollow ball, the wall of which consists of a single stratum of cells (Figure 1.38 A to C). This layer is the blastoderm, the simple epithelium from the cells of which all the tissues of the body proceed. These important early embryonic processes take place so quickly in the Amphioxus that four or five hours after fecundation, or about midnight, the spherical blastula is completed. A pit-like depression is then formed at the vegetal pole of it, and in consequence of this the hollow sphere doubles on itself (Figure 1.38 D). This pit becomes deeper and deeper (Figure 1.38 E and F); at last the invagination (or doubling) is complete, and the inner or folded part of the blastula-wall lies on the inside of the outer wall. We thus get a hollow hemisphere, the thin wall of which is made up of two layers of cells (Figure 1.38 E). From hemispherical the body soon becomes almost spherical once more, and then oval, the internal cavity enlarging considerably and its mouth growing narrower (Figure 2.213). The form which the Amphioxus-embryo has thus reached is a real "cup-larva" or gastrula, of the original simple type that we have previously described as the "bell-gastrula" or archigastrula (Figures 1.29 to 1.35). As in all the other animals that form an archigastrula, the whole body is nothing but a simple gastric sac or stomach; its internal cavity is the primitive gut (progaster or archenteron, Figure 1.38 g, 1.35 d), and its aperture the primitive mouth (prostoma or blastoporus, o). The wall is at once gut-wall and body-wall. It is composed of two simple |
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