_F1 grossulariata_[female] it produced all the [male]'s _grossulariata_
and all the [female]'s _lacticolor_. Bateson's explanation is that the
female, according to the Mendelian theory of sex, is heterozygous in sex,
the male homozygous and recessive, and that _lacticolor_ is linked with
the female sex-character, _grossulariata_ being repelled by that
character. Thus we have, the _lacticolor_ character being recessive,

lact. male, LL male male x F, gross. female, GL female male
Gametes L male + L male x G male + L female
_____________________|______________________
| |
GL male male LL male female
gross. male lact. female

It will be seen that although in the progeny of this mating all the
_grossulariata_ were males and all the _lacticolor_ females, yet this case
is by no means similar to that of sexual dimorphism in which the
characters are normally always confined to the same sex. For the
_lacticolor_ character in the parent was in the male, while in the
offspring it was in the female. We cannot say here that in the theoretical
factors which are supposed to represent what happens, the _lacticolor_
character is coupled with the female sex-factor, for we find it with the
male sex-character in the _lacticolor_ [male]. It is so coupled only in
the heterozygous _grossulariata_ [female], and at the same time the
_grossulariata_ character is repelled.

According to Doncaster [Footnote: _Determination of Sex_, Camb. Univ.
Press, 1914.] sex-limited, or as it is now proposed to call it sex-linked,
transmission in this case means that the female _grossulariata_ transmits
the character to all her male offspring and to none of the female, while a